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, GRX1-roGFP2 or TRaQ-G) enable real-time, organelle-specific redox readouts but primarily report redox potential rather than absolute GSH levels, whereas fractionation coupled with mass spectrometry provides absolute GSH/GSSG quantification but remains static and susceptible to lysis-associated artifacts

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Resulting segmentation maps, combined with ( X , Y ) probe coordinates, were used for clustering and graph-based analysis (Extended Data Fig