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Acknowledgements This work was supported by National Natural Science Foundation of China (Grant number: 30400077), University of Gdask (task grant no

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Since in some cases gshA and gshB fuse to form a single gene gshF and sequence divergence is very high for both GshA and GshB [3], the results of GshA, GshB and GshF were manually checked for the position of alignment in the gene and length of the hit